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24, chemin de Borde Rouge –Auzeville – CS52627
31326 Castanet Tolosan CEDEX - France

Dernière mise à jour : Mai 2018

Menu Logo Principal AgroParisTech Université Paris-Saclay


GABI : Génétique Animale et Biologie IntégrativeUnité Mixte de Recherche INRA - AgroParisTech

Didier BOICHARD, Senior Research Scientist

Genomic selection; genetic Improvement of Cattle; genetic evaluation; association analysis ; milk composition ; disease resistance; genetic defects
Fond Blanc2

INRAE, UMR 1313 Animal Genetics and Integrative Biology
Domaine de Vilvert, Bat 211, 78352 Jouy en Josas, France
Tel : +33 (0) 1 34 65 21 81

Email : didier.boichard(at)

Research Team: Bovine Genetics and Genomics

CV :

  • Agronomic Engineer from the Institut National Agronomique Paris Grignon, 1982
  • MSc in Quantitative Genetics, Paris VI, 1985
  • PhD from INA Paris Grignon (presently AgroParisTech), 1987
  • Junior Research Scientist (1987) then Senior Research Scientist (1996) at INRAE
  • Head of the Animal Genetics Division of INRAE from 2002 to 2009
  • Chairman of Labogena from 2004 to 2013
  • Member of the Veterinary Academy of France
  • Gold medal of the Agriculture Academy of France (2015)
  • G2B team recieved the Laurier for Impact award from INRAE in 2016

Fields of Research :

Genomic selection ; Genetic Improvement of cattle; Quantitative Genetics; Genetic evaluation; QTL mapping; Genetic determinism of milk composition; Genetic determinism of fertility; Genetic determinism of mastitis resistance; Genetic defects.

Other Activites :

  • Coordination : Head of the G2B research team ; co-leader of the UMT 3G (2011-16) then of the UMT eBIS (2017-22)
  • Expertise : Member of the National Committee on Animal Selection ; Member of the executive committee of France Génétique Elevage
  • Publication : Co-Editor in Chief of Genetics Selection Evolution

Selected Publications :

Boichard D. 2020. Nouvelles modalités de diffusion de la génétique. In “Génétique des animaux d'élevage : Diversité et adaptation dans un monde changeant”, Quae, E Verrier, D Milan, C Rogel-Gaillard Eds, chapitre 12, 193-203.

Sanchez M.P., Fritz S., Patry C., Delacroix-Buchet A., Boichard D. 2020. Milk protein variant and haplotype frequencies estimated from genotypes of more than one million bulls and cows of twelve French cattle breeds. J Dairy Sci, 103, 9124-9141.

Sanchez M.P., Guatteo R., Davergne A., Saout J., Grohs C., Deloche M.C., Taussat S., Fritz S., Boussaha M., Blanquefort P., Delafosse A., Joly A., Schibler L., Fourichon C., Boichard D. 2020. Identification of the ABCC4, IER3 and CBFA2T2 candidate genes for resistance to paratuberculosis in Holstein and Normande dairy cattle from a sequence-based GWAS. Genet Sel Evol, 52,14.

Zhang J., Kadri N., Mullaart E., Spelman R., Fritz S., Boichard D., Charlier C., Georges M., Druet T. 2020. Genetic architecture of individual variation in recombination rate on the X-chromosome in cattle, Heredity, early online.

Van Den Berg I., Xiang R., Jenko J., Pausch H., Boussaha M., Tribout T., Gjuvsland A.B., Boichard D., Nordbø Ø., Sanchez M-P., Goddard M.E. 2020. Large scale multi breed meta-analysis for fat and protein percentage using imputed sequence variant genotypes in 94,321 cattle from eight dairy breeds. Genet Sel Evol, 52, 37.

Liu A., Lund M.S., Boichard D., Karaman E., Guldbrandtsen B., Fritz S., Aamand G.P., Nielsen U.S., Sahana G., Wang Y., Su G. 2020. Weighted single-step genomic best linear unbiased prediction integrating the variants selected from sequencing data by association and bioinformatics analyses. Genet Sel Evol, 52, 48.

Liu A., Lund M.S., Boichard D., Mao X., Karaman E., Fritz S., Pedersen Aamand G., Wang Y., Su G. 2020. Imputation for sequencing variants preselected to a customized low-density chip. Sci Rep, 10:9524.

Boichard D. 2019. Development of dairy breeding programs. Book “Advances of breeding in dairy cattle”, Burleigh-Dodds, Chapitre 23, 16p. 

Hozé C., Escouflaire C., Mesbah-Uddin M., Barbat A., Boussaha M., Deloche M.C., Boichard D., Fritz S., Capitan A. 2020. Short Communication: A splice site mutation in CENPU is associated with recessive embryonic lethality in Holstein cattle. J Dairy Sci, 103:607–612.

Liu A., Lund M.S., Boichard D., Karaman E., Fritz S., Aamand G.P., Nielsen U.S., Wang Y., Su G. 2020. Improvement of genomic prediction by additional single nucleotide polymorphisms selected from whole genome sequencing data. Heredity, 124, 37-49.

Sanchez M.P., Wolf V., Laithier C., El Jabri M., Beuvier E., Rolet-Repecaud O., Gaudilliere N., Minery S., Ramayo-Caldas Y., Tribout T., Michenet A., Boussaha M., Taussat S., Fritz S., Delacroix-Buchet A., Grosperrin P., Brochard M., Boichard D. 2019. Analyse génétique de la fromageabilité du lait de vache prédite par spectrométrie moyen infrarouge en race Montbéliarde. INRA Productions Animales, 32, 379-398.

Dezetter C., Boichard D., Bareille N., Grimard B., Le Mezec P., Ducrocq V. 2019. Le croisement entre races bovines laitières : Quel intérêt pour des ateliers en race pure Prim’Holstein ? INRA Productions Animales, 32, 359-378.

Sanchez M.P., Ramayo-Caldas Y., Wolf V., Laithier C., El Jabri M., Michenet A., Boussaha M., Taussat S., Fritz S., Delacroix-Buchet A., Brochard M., Boichard D. 2019. Sequence-based GWAS, network and pathway analyses reveal genes co-associated with milk cheese making properties and milk composition in Montbéliarde cows. Genet Sel Evol, 51, 34.

El Jabri M., Sanchez M.P., Trossat P., Laithier C., Wolf V., Grosperrin P., Beuvier E., Rolet-Répécaud O., Gavoye S., Gauzere Y., Belysheva O., Notz E., Boichard D., Delacroix-Buchet A. 2019. Comparison of Bayesian and PLS regression methods for mid-infrared prediction of cheese-making properties in Montbéliarde cows. J Dairy Sci., 102:6943–6958.

Uddin M.M., Hoze C., Michot P., Barbat A., Lefebvre R., Boussaha M., Sahana G., Fritz S., Boichard D., Capitan A. 2019. A missense mutation (p.Tyr452Cys) on CAD gene compromises reproductive success in dairy cattle. J Dairy Sci., 102:6340–6356.

Mesbah-Uddin M, Guldbrandtsen B, Lund MS, Boichard D, Sahana G. 2019. Joint imputation of whole-genome sequence variants and large chromosomal deletions in cattle. J Dairy Sci. 102:11193-11206

Navarro-Gonzalez N, Fourichon C, Blanquefort P, Delafosse A, Joly A, Ngwa-Mbot D, Biet F, Boichard D, Schibler L, Journaux L, Meens E, Guatteo R. 2019. Longitudinal Study of Mycobacterium avium subsp paratuberculosis Shedding Patterns and Concurrent Serological Patterns in Naturally Infected Dairy Cattle. J Dairy Sci. 102, 9117-9137.

Bérodier M., Brochard M., Boichard D., Dezetter C., Bareille N., Ducrocq V. 2019. Use of sexed semen and female genotyping impacts genetic and economic outcomes of Montbéliarde dairy herds depending on the farming system considered. J Dairy Sci, 102:10073–10087.

Sellier P, Boichard D, Verrier E. 2019. La génétique animale à l‘Inra : soixante ans d'une histoire scientifique en prise avec le monde de la sélection et riche en rebondissements technologiques. Histoire Recherche Contemporaine. 8, 86-98.

Boichard D, Mattalia S, Fritz S. 2018. La sélection dans l’espèce bovine et ses perspectives. Bull Acad Vet. 170, 215-220.

Sanchez M.P., El Jabri M., Minery S., Wolf V., Beuvier E., Laithier C., Delacroix-Buchet A., Brochard M., Boichard D. 2018. Genetic parameters for cheese-making properties and milk composition predicted from mid-infrared spectra in a large dataset of Montbéliarde cows. J Dairy Sci, 101, 10048–10061.

Sanchez M.P., Wolf V., El Jabri M., Beuvier E., Rolet-Repecaud O., Gauzere Y., Minery S., Brochard M., Michenet A., Taussat S., Barbat-Leterrier A., Delacroix-Buchet A., Laithier C., Fritz S., Boichard D. 2018. Short-communication: Confirmation of candidate causative variants on milk composition and cheese-making properties in Montbéliarde cows. J Dairy Sci, 101, 10076–10081.

Marete A., Guldbrandtsen B., Lund M.S., Sahana G., Boichard D. 2018. A meta-analysis including pre-selected sequence variants associated with seven traits in three French dairy cattle populations. Frontiers in Genetics, 9, 522.

Duchesne A., Vaiman A., Frah M., Floriot S., Rodriguez S., Desmazieres A., Fritz S., Beauvallet C., Albaric O., Venot E., Bertaud M., Guatteo R., Esquerre D., Brice A., Vilotte J.L., Darios F., Stevanin G., Boichard D., El Hachimi K.H. 2018. Progressive ataxia of Charolais cattle highlights a role of KIF1C in sustainable myelination. Plos Genetics, 14: e1007550.

Rainard P., Foucras G., Boichard D., Rupp R. 2018. Invited review: Low milk somatic cell count and susceptibility to mastitis. J Dairy Sci., 101, 6703-6714.

Rainard P., Foucras G., Boichard D., Rupp R. 2018. Faibles concentrations cellulaires du lait et sensibilité aux mammites des ruminants laitiers. INRA Prod Anim, 31, 365-376.

Perrier J.P., Sellem E., Prezelin A., Gasselin M., Jouneau L., Piumi F., Al Adhami H., Weber M., Fritz  S., Boichard D., Le Danvic C., Schibler L., Jammes H., Kiefer H. 2017. A multi-scale analysis of bull sperm methylome revealed both species peculiarities and conserved tissue-specific features. BMC Genomics, 19, 404.

Fritz S., Hoze C., Rebours E., Barbat A., Bizard M., Chamberlain A., Escouflaire C., Vander Jagt C., Boussaha M., Grohs C., Allais-Bonnet A., Philippe M., Vallee A., Amigues Y., Hayes B.J., Boichard D., Capitan A. 2018. An initiator codon mutation in SDE2 causes recessive embryonic lethality in Holstein cattle. J Dairy Sci, 101, 6220-6231.

Marete A.G., Boichard D., Lund M.S., Ramayo-Caldas Y. 2018. A system-based analysis of the genetic determinism of udder conformation and health traits across three French Dairy Breeds. Plos One, 13, e0199931.

Marete A.G., Sahana G., Fritz S., Lefebvre R., Barbat-Leterrier A., Lund M.S., Guldbrandtsen B., Boichard D. 2018. Genome-wide association study for milking speed in French Holstein cows. J Dairy Sci., 101, 6205-6219.

Bouwman A.C., Daetwyler H.D., Chamberlain A.J., Hurtado Ponce C., Sargolzaei M., Schenkel F.S., Sahana G., Govignon-Gion A., Boitard S., Dolezal M., Pausch H., Brøndum R.F., Bowman P.J., Thomsen B., Guldbrandtsen B., Lund M.S., Servin B., Garrick D.J., Reecy J., Vilkki J., Bagnato A., Wang M., Hoff J.L., Schnabel R.D., Taylor J.F., Vinkhuyzen A.A.E., Panitz F., Bendixen C., Holm L.E., Gredler B., Hozé C., Boussaha M., Sanchez M.P., Rocha D., Capitan A., Tribout T., Barbat A., Croiseau P., Drögemüller C., Jagannathan V., Vander Jagt C., Crowley J.J., Intergenomics Consortium, Bieber A., Purfield D.C., Berry D.P., Emmerling R., Götz K.U., Van Tassell C.P, Fries R., Stothard P., Veerkamp R.F., Boichard D., Goddard M.E., Hayes B.J.  2018. Meta-analysis of genome wide association studies for the stature of cattle reveals common genes that regulate size in mammals. Nature Genetics, 50, 362-367, DOI:10.1038/s41588-018-0056-5.

Charton C., Guinard-Flament J, Lefebvre R, Barbey S., Gallard Y, Boichard D, Larroque H. 2017. Genetic determinism of milk production traits in response to a three-week once-daily milking period in crossbred Holstein × Normande dairy cows. J Dairy Sci., 101, 2235–2247.

Mesbah-Uddin M., Guldbrandtsen B., Iso-Touru T., Vilkki J., De Koning D.J., Boichard D., Lund M.S., Sahana G. 2018. Genome-wide mapping of large deletions and their population-genetic properties in dairy cattle. DNA Research, 25, 49-59.

Sanchez M.P., Govignon-Gion A., Croiseau P., Fritz S., Hoze C., Miranda G., Martin P., Barbat-Leterrier A., Brochard M., Boussaha M., Boichard D. 2017. Within-breed and multi-breed GWAS on imputed whole genome sequence variants reveal candidate mutations affecting milk protein composition in dairy cattle. Genetics Selection Evolution, 49: 68.

Bourneuf E., Otz P., Pausch H., Jagannathan V., Michot P., Grohs C., Piton G., Ammermüller S., Deloche M.C., Fritz S., Leclerc H., Péchoux C., Boukadiri A., Saintilan R., Créchet F., Mosca M., Segelke D., Guillaume F., Bouet S., Baur A., Vasilescu A., Genestout L., Thomas A., Allais-Bonnet A., Rocha D., Colle M.A., Klopp C., Esquerré D., Wurmser C., Flisikowski K., Schwarzenbacher H., Burgstaller J., Brügmann M., Dietschi E., Huth N., Freick M., Barbey S., Fayolle G., Danchin-Burge C., Schibler L., Bed’hom B., Hayes B.J., Daetwyler H.D., Fries R., Boichard D., Pin D., Drögemüller C., Capitan A. 2016. Rapid Discovery of De Novo Deleterious Mutations in Cattle Using Genome Sequence Data: Enhancing the Value of Farm Animals as Model Species. Scientific Reports, 7: 11466.

Michot P., Fritz S., Barbat A., Boussaha M., Deloche M.C., Grohs C., Hoze C., Le Berre L., Le Bourhis D., Desnoes O., Salvetti P., Schibler L., Boichard D., Capitan A. A missense mutation in PFAS is likely causal for embryonic lethality associated with the MH1 haplotype in Montbeliarde dairy cattle. J Dairy Sci, 100, 8176–8187.

Duchesne A., Vaiman A., Castille J., Beauvallet C., Gaignard P., Floriot S., Rodriguez S., Vilotte M., Boulanger L., Passet B., Albaric O., Guillaume F., Boukadiri A., Richard L., Bertaud M., Timsit E., Guatteo R., Jaffrézic F., Calvel P., Mahla R., Esquerré D., Péchoux C., Liu S., Vallat J.M., Boichard D., Slama A., Vilotte J.L. 2017. Bovine and murine models highlight novel roles for SLC25A46 in mitochondrial dynamics and metabolism, with implications for human and animal health. Plos Genet., 13, e1006597.

Fang Z.H., Bovenhuis H., Delacroix-Buchet A., Miranda G., Boichard D., Visker M.H.P.W., Martin P. 2017. Genetic and non-genetic factors contributing to differences in alpha-s-casein phosphorylation isoforms and other major milk proteins. J Dairy Sci., 100, 5564-5577.

Letaief R, Rebours E, Grohs C, Meersseman C, Fritz S, Trouilh L, Esquerré D, Barbieri J, Klopp C, Philippe R, Blanquet V, Boichard D, Rocha D, Boussaha M. 2017. Identification of copy number variation in French dairy and beef breeds using next-generation sequencing. Genet Sel Evol. 49, 77.

Sanchez M.P., Govignon-Gion A., Ferrand M., Gele M., Pourchet D., Miranda G., Martin P., Brochard M., Boichard D. 2017. Short-communication: Genetic parameters for milk protein composition in the French Montbéliarde, Normande and Holstein dairy cattle breeds. Journal of Dairy Science, 100, 6371-6375.

Boichard D, Ducrocq V, Croiseau P, Fritz S. 2016. Genomic selection in domestic animals: Principles, applications and perspectives. CR Biol. 339, 274-277.

Boichard D. , Grohs C. , Danchin C., Capitan A. 2016. Anomalies génétiques : définition, principes et concepts. INRA Productions Animales, 29, 297-306.

Grohs C, Duchesne A, Floriot S, Deloche MC, Boichard D, Ducos A, Danchin-Burge C. 2016. L’Observatoire National des Anomalies Bovines, son action et ses résultats pour une aide efficace à la gestion des anomalies génétiques. Inra Prod Anim. 29, 307-318.

Duchesne A, Grohs C, Michot P, Bertaud M, Boichard D, Floriot S, Capitan A. 2016. Du phénotype à la mutation causale : le cas des anomalies récessives bovines. Inra Prod Anim. 29, 319-328.

Fritz S.,  Michot P, Hoze C., Grohs C, Barbat A., Boussaha M., Boichard D, Capitan A. 2016. Anticiper l'émergence d'anomalies génétiques grâce aux données génomiques. INRA Productions Animales, 29, 339-350.

Boichard D, Grohs C, Michot P, Danchin C, Capitan A, Genestout L, Barbier S., Fritz S. 2016. Prise en compte des anomalies génétiques en sélection. INRA Productions Animales, 29, 351-358.

Boussaha M., Michot P., Letaief R., Hoze C., Fritz S., Grohs C., Esquerre D., Duchesne A., Philippe R., Blanquet V., Phocas F., Floriot S., Rocha D., Klopp C., Capitan A., Boichard D. 2016. Construction of a large collection of small genome variations in French dairy and beef breeds using whole genome sequences. Genetics Selection Evolution, 48, 87.

Kadri N.K., Harland C., Faux P., Cambisano N., Karim L., Coppieters W., Fritz S., Mullaart E., Baurain D., Boichard D., Spelman R., Charlier C., Georges M., Druet T. 2016. Sequence-based association analysis identifies coding and non coding variants in HFM1, MLH3, MSH4, MSH5, RNF212 and RNF212B with large effects on male and female recombination rate in cattle. Genome Research, 26, 1323-1332.

Van Den Berg I., Boichard D., Lund M.S. 2016. Sequence variants selected from a multi breed GWAS can improve the reliability of genomic prediction in dairy cattle. Genetics Selection Evolution, 48, 83.

Van Den Berg I., Boichard D., Lund M.S. 2016. Comparing power and precision of within and multi breed genome wide association studies of production traits using whole genome sequence data for five French and Danish dairy cattle breeds. Journal of Dairy Science, 99, 8932–8945.

Van Den Berg  I., Boichard D., Guldbrandtsen B., Lund M.S. 2016. Use of sequence variants to improve accuracy of genomic prediction in dairy cattle breeds. G3, 6, 2553-2561.

Michot P., Chahory S., Marete A., Grohs C., Dagios D., Donzel E., Aboukadiri A., Deloche M.C., Allais-Bonnet A., Chambrial M., Barbey S., Boussaha M., Danchin-Burge C., Fritz S., Boichard D., Capitan A. 2016. A reverse genetic approach identifies an ancient frameshift mutation in RP1 causing recessive progressive retinal degeneration in European cattle breeds. Genetics Selection Evolution, 48, 56.

Sanchez M.P. , Govignon-Gion A., Ferrand M., Gele M., Pourchet D., Amigues Y., Fritz S., Boussaha M., Capitan A., Rocha D., Miranda G., Martin P., Brochard M., Boichard D. 2016. Whole genome scan to detect SNP associated to milk protein composition in three French dairy cattle breeds. Journal of Dairy Science, 99, 8203–8215

Colleau J. J., Fritz S., Guillaume F., Baur A., Dupassieux D., Boscher M.Y., Journaux L., Eggen A., Boichard D. 2015. Simulation des potentialités de la sélection génomique chez les bovins laitiers. INRA Productions Animales, 28, 251-258.

Boussaha M., Esquerre D., Barbieri J., Djari A., Pinton A., Letaief R., Salin G., Escudie F., Roulet A., Fritz S., Samson F., Grohs C., Bernard M., Klopp C., Boichard D., Rocha D. 2015. Genome-wide study of structural variants in bovine Holstein, Montbeliarde and Normande dairy breeds. Plos One, 10: e0135931.

Dezetter C., Leclerc H., Mattalia S., Barbat A., Boichard D., Ducrocq V. 2015. Inbreeding and Crossbreeding Parameters for Production and Fertility Traits in Holstein, Montbéliarde, and Normande Cows. Journal of Dairy Science, 98, 4904–4913.

Brøndum R.F, Su G., Janss L., Sahana G., Guldbrandtsen B., Boichard D., Lund M.S. 2015. QTL markers derived from whole genome sequence data increases the reliability of genomic prediction. J Dairy Sci., 98, 4107-4116.

Floriot S., Vesque C., Rodriguez S., Bourgain-Guglielmetti F., Karaiskou A., Gautier M., Duchesne A., Barbey S., Fritz S., Vasilescu A., Bertaud M., Moudjou M., Halliez S., Cormier-Daire V., El Hokayem J., Nigg E.A., Manciaux L., Guatteo R., Cesbron N., Toutirais G., Eggen A., Schneider-Maunoury S., Boichard D., Sobczak-Thépot J., Schibler L.  2015. C-Nap1 mutation affects centriole cohesion and is associated with a Seckel-like syndrome in cattle. Nature Communications, 6, 6894.

Michot P., Fantini O., Braques R., Allais-Bonnet A., Saintilan R., Grohs C., Barbieri J., Genestout L., Danchin C., Gourreau J.M., Boichard D., Pin D., Capitan A. 2015. Whole-genome sequencing identifies a homozygous deletion encompassing exons 17 to 22 of the Integrin Beta 4 Gene in a Charolais calf with Junctional Epidermolysis Bullosa. Genetics Selection Evolution, 47, 37.

Baumung R., Farkas J., Boichard D., Meszaros G., Sölkner J., Curik I. 2015. GRAIN: a computer program to calculate ancestral and partial inbreeding coefficients using a gene dropping approach. Journal of Animal Breeding and Genetics, 132, 100-108.

Boichard D., Ducrocq V., Fritz S. 2015. Sustainable dairy cattle selection in the genomic era. Journal of Animal Breeding and Genetics, 132, 135-143.

Capitan A., Michot P., Baur A., Saintilan R., Hoze C., Valour D., Guillaume F., Boichon D., Barbat A., Boichard D., Schibler L., Fritz S. 2015. Genetic tools to improve reproduction traits in dairy cattle. Reprod. Fert. Dev, 27, 14-21

Boichard D. 2014. Quel futur pour l’amélioration génétique chez les espèces domestiques ? Bulletin de l’Académie Vétérinaire de France,167, 103-108.

Boichard D., Govignon-Gion A., Larroque H., Maroteau C., Palhiere I., Tosser-Klopp G., Rupp R., Sanchez M.P., Brochard M. 2014. Déterminisme génétique de la composition en acides gras et protéines du lait des ruminants. INRA Productions Animales, 27, 283-298.

Daetwyler H.D., Capitan A., Pausch H., Stothard P.,  Van Binsbergen R., Brøndum R.F., Liao X., Djari A., Rodriguez S.C., Grohs C., Esquerré D., Bouchez O., Rossignol M.N., Klopp C., Rocha D., Fritz S., Eggen A., Bowman P., Coote D., Chamberlain A.J., Anderson C., Van Tassell C.P., Hulsegge I., Goddard M.E., Guldbrandtsen B., Lund M.S., Veerkamp R.F., Boichard D., Fries R.,  Hayes B.J. 2014. Whole-genome sequencing of 234 bulls facilitates mapping of monogenic and complex traits in cattle. Nature Genetics, 46, 858-867.

Hozé C., Fritz S., Phocas F., Boichard D., Ducrocq V., Croiseau P. 2014. Efficiency of multi-breed genomic selection for dairy cattle breeds with different sizes of reference population. Journal of Dairy Science 97, 3918-3929.

Labatut J., Astruc J.M., Barillet F., Boichard D., Ducrocq V., Griffon L., Lagriffoul G. 2014. Implications organisationnelles de la sélection génomique: analyses et accompagnement. INRA Productions Animales, 27, 303-316.

Rocha D., Billerey C., Boichard D., Boussaha M. 2014. Identification of the putative ancestral allele of bovine single nucleotide polymorphisms. Journal of Animal Breeding and Genetics, 131, 483-486.

Van Den Berg I., Fritz I., Rodriguez S., Rocha D., Boussaha M., Lund M.S., Boichard D. 2014. Concordance analysis for QTL detection in dairy cattle: a case study of leg morphology. Genetics Selection Evolution, 46, 31.

Allais-Bonnet A., Grohs C., Medugorac I., Krebs S., Djari A., Graf A., Fritz S., Seichter D., Baur A., Bouet S., Wahlberg P., Esquerre D., Hoze C., Boussaha M., Weiss B., Thepot D., Venot E., Fouilloux M.N., Rossignol M.N., Van Marle-Köster E., Hreiðarsdottir G.E., Barbey S., Dozias D., Cobo E., Reverse P., Catros O., Marchand J.L., Soulas P., Roy P., Marquant-Leguienne B., Lebourhis D., Pannetier M., Phocas F., Klopp C.,  Rocha D., Fouchet M., Journaux L., Bernard-Capel C., Eggen A., Förster M., Gallard Y., Boichard D., Pailhoux E., Capitan A. 2013. Novel insights into the bovine polled phenotype and horn ontogenesis in Bovidae. Plos One 8, e63512.

Boichard D., 2013. La sélection génomique : une opportunité pour l’amélioration de la santé des animaux d’élevage. Bulletin de l’Académie Vétérinaire de France, 166, 25-31.

Brochard M., Boichard D., Ducrocq V., Fritz S. 2013. La sélection pour des vaches et une production laitière plus durables : acquis de la génétique et opportunités offertes par la sélection génomique. INRA Prod. Anim., 26, 139-150.

Fritz S., Capitan A., Djari A., Rodriguez S.C., Barbat A., Baur A., Grohs C., Weiss B., Boussaha M., Esquerre D., Klopp C., Rocha D., Boichard D. 2013. Detection of haplotypes associated with prenatal death in dairy cattle and identification of deleterious mutations in GART, SHBG, and SLC37A2. Plos One, 8, e65550.

Hoze C., Fouilloux M.N., Venot E., Guillaume F., Dassonneville R., Fritz S., Ducrocq V., Phocas F., Boichard D., Croiseau P. 2013. High density marker imputation efficiency in 16 French cattle breeds. Genet. Sel. Evol, 45, 33.

Van Den Berg I., Fritz S., Boichard D. 2013. QTL fine mapping with Bayesian C(π): a simulation study. Genet. Sel Evol., 45, 19.

Boichard D., Brochard M. 2012.  New phenotypes for new breeding goals in dairy cattle. Animal, 6, 544-550.

Boichard D., Guillaume F., Baur A., Croiseau P., Rossignol M.N., Boscher M.Y., Druet T., Genestout L., Colleau J.J., Journaux L., Ducrocq V., Fritz S. 2012. Genomic Selection in French Dairy Cattle. Animal Production Science, 52, 115-120.

Boichard D., Chung H., Dassonneville R., David X., Eggen A., Fritz S., Gietzen K.J., Hayes B.J., Lawley C.T., Sonstegard T.S., Van Tassell C.P., Vanraden P.M., Viaud K., Wiggans G.R. 2012. Design of a Bovine Low-Density SNP Array Optimized for Imputation. PLoS ONE, 7: e34130.

Capitan A., Bonnet A., Pinton A., Marquant-Le Guienne B., Le Bourhis D., Grohs C., Bouet S., Clement L., Salas-Cortes L., Venot E., Chaffaux S., Weiss B., Delpeuch A., Noe G., Rossignol M.N., Barbey S., Dozias D., Cobo E., Barasc H., Auguste A., Pannetier M., Deloche M.C., Lhuilier E., Bouchez O., Esquerre D., Salin G., Klopp C., Donnadieu C., Chantry-Darmon C., Hayes H., Gallard Y, Ponsart C., Boichard D., Pailhoux E. 2012. A 3.7 Mb deletion encompassing ZEB2 causes a novel Polled and Multisystemic Syndrome in the progeny of a somatic mosaic bull. Plos One, 7: e49084.

Dassonneville R., Baur A., Fritz S., Boichard D., Ducrocq V. 2012. Inclusion of cows records in genomic evaluations and its impact on bias due to preferential treatment. Genet. Sel. Evol., 44:40.

Dassonneville R., Fritz S., Ducrocq V., Boichard D. 2012. Imputation performances of three low density marker panels in beef and dairy cattle. Journal of Dairy Science, 95: 4136–4140.

Dayo G.K., Gautier M., Berthier D., Poivey J.P., Sidibe I., Bengaly Z., Eggen A., Boichard D., Thevenon S. 2012. Association studies in QTL regions linked to bovine trypanotolerance in a West African Crossbred Population. Animal Genetics, 43, 123-132.

Hou Y., Bickhart D.M., Hvinden M.L., Li C., Song J., Boichard D., Fritz S., Eggen A., Denise S.,  Wiggans G.R., Sonstegard T.S., Van Tassell C.P., Liu G.E. 2012. Fine mapping of copy number variations on two cattle genome assemblies using high density SNP array. BMC Genomics, 13, 376.

Ytournel F., Teyssedre S., Roldan D., Erbe M., Simianer H., Boichard D., Gilbert H., Druet T., Legarra A. 2012. LDSO: A program to simulate pedigrees and molecular information under various evolutionary forces. Journal of Animal Breeding and Genetics, 5, 417-421.

Croiseau P., Legarra A., Guillaume F., Fritz S., Baur A., Boichard D., Ducrocq V. 2011. Fine tuning genomic evaluations in dairy cattle through SNP pre-selection with the Elastic-Net algorithm. Genetics Research, 93, 409-417.

Guillaume F., Boichard D., Ducrocq V., Fritz S. 2011. Utilisation de la sélection génomique chez les bovins laitiers. INRA Prod. Anim., 24,363-368.

Barbat A., Le Mezec P., Ducrocq V., Mattalia S., Fritz S., Boichard D., Ponsart C., Humblot P. 2010. Female Fertility in French Dairy Breeds: Current Situation and Strategies for Improvement. Journal of Reproduction and Development, 56, S15-S21.

Bidanel J.P., Boichard D., Chevalet C. 2008. De la génétique à la génomique. INRA Productions Animales, 21, 15-32.

Druet T., Fritz S., Boussaha M., Ben Jemaa S., Derbala D., Zele D., Lechner D., Charon C., Boichard D., Gut I.G., Eggen A., Gautier M. 2008. Fine-mapping of QTLs affecting female fertility in dairy cattle on BTA03 using a dense SNP map. Genetics, 178, 2227-2235.

Guillaume F., Fritz S., Boichard D., Druet T. 2008. Estimation by simulation of the efficiency of the French marker-Assisted selection program in dairy cattle. Genetics Selection Evolution, 40, 91-102.

Gautier M, Fritz S., Capitan A., Eggen A., Boichard D., Druet T. 2007. Characterization of the K232A and VNTR polymorphisms in the DGAT1 gene in French dairy cattle. J. Dairy Sci, 90: 2980-2988.

Gautier M., Faraut T., Moazami-Goudarzi K., Navratil V., Foglio M., Grohs C., Boland A., Garnier J.G., Boichard D., Lathrop G.M., Gut I.G., Eggen A. 2007. Genetic and haplotypic structure in 14 European and African cattle breeds. Genetics, 177, 1059-1070.

Druet T., Fritz S., Boichard D., Colleau J.J. 2006. Estimation of genetic parameters for quantitative trait loci for dairy traits in the French Holstein population. Journal of Dairy Science, 89, 4070-4076.

Barnouin J., Chassagne M., Bazin S., Boichard D., 2004. Management and hygiene practices form questionnaire surveys in herds with very low somatic cell score through the ‘Objective Zero Mastitis’ national programme in France J Dairy Sci, 87:3989-3999.

Bennewitz J, Reinsch N., Guiard V., Fritz S., Thomsen H., Looft C., Kühn C., Schwerin M., Weimann C., Erhardt G., Reinhardt F., Reents R., Boichard D., Kalm E. 2004. Multiple Quantitative Trait Loci Mapping With Cofactors and Application of Alternative Variants of the False Discovery Rate in an Enlarged Granddaughter Design. Genetics 168: 1019–1027.

Schibler L., Roig A., Mahe M.F., Save J.C., Gautier M., Taourit S., Boichard D., Eggen A., Cribiu E.P.. 2004. A first generation bovine BAC-based physical map. Genet. Sel. Evol. 36, 105-122.

Boichard D., Grohs C., Bourgeois F., Cerqueira F., Faugeras R., Neau A., Rupp R., Amigues Y., Boscher M.Y., Leveziel H., 2003. Detection of genes influencing economic traits in three French dairy cattle breeds. Genet. Sel. Evol. 35, 77-101.

Druet T., Jaffrezic F., Boichard D., Ducrocq V., 2003. Estimation of genetic parameters for first parity lactation curves of French Holstein cows. J. Dairy Sci., 86: 2480 - 2490.

Rupp R., Boichard D., 2003. Genetics of resistance to mastitis in dairy cattle. Vet. Res., 34, 671-688.

Rupp R., Boichard D. 2000. Relationship of early first lactation somatic cell count with risk of subsequent first clinical mastitis. Livest. Prod. Sci., 62, 169-180

Rupp R., Beaudeau F. Boichard D., 2000. Relationship between milk somatic cell counts in the first lactation and clinical mastitis in the second lactation of French Holstein cows. Prev.Vet. Med., 46, 99-111.

Rupp R., Boichard D. 1999. Genetic parameters for clinical mastitis, somatic cell score, production, udder type traits, and milking ease in first lactation Holsteins. J. Dairy Sci., 82, 2198-2204.

Robert-Granie C., Bonaiti B., Boichard D., Barbat A. 1999. Accounting for variance heterogeneity in French dairy cattle genetic evaluation. Livest. Prod. Sci., 60, 343-357

Zhang Q., Boichard D., Bishop M., Hoeschele I., Ernst C., Doud L., Eggen A., Jurgella G., Murkve B., Pfister-Genskow M., Thorbahn D., Grignola Fe, Uimari P., Thaller G. 1998. Mapping quantitative trait loci for milk production and health of dairy cattle in a large outbred pedigree. Genetics, 149, 1959-1973

Boichard D., Maignel L., Verrier E. 1997. The value of using probabilities of gene origin to measure genetic variability in a population. Genet. Sel. Evol., 29, 5-23.

Boichard D., Bonaiti B., Barbat A., Mattalia S. 1995. Three methods to validate the estimation of genetic trend for dairy cattle. J. Dairy Sci., 78, 431-437.

Boichard D., Manfredi E. 1994. Genetic analysis of conception rate in French Holstein dairy cattle. Acta Agric. Scand., Sect. A, 44, 138-145.

Foulley J.L., Hanocq E., Boichard D. 1992. A criterion for measuring the degree of connectedness in linear models of genetic evaluation. Genet. Sel. Evol., 24, 315-330.

Boichard D. 1990. Estimation of the economic value of conception rate in dairy cattle. Livest. Prod. Sci., 24, 187-204.

Boichard D, Bonaiti B. 1987. Genetic parameters for first lactation dairy traits in Friesian, Montbéliarde and Normande breeds. Genet. Sel. Evol., 19, 337-350.